/* ==========================================================================
   鳢科图谱 · Channidae Atlas
   ========================================================================== */

:root{
  --paper:#f3eee2;
  --paper-2:#faf6ec;
  --paper-3:#ece4d2;
  --ink:#18211c;
  --ink-2:#333d36;
  --muted:#6d7469;
  --muted-2:#8b9186;
  --line:#d9cfba;
  --line-soft:#e7dfcd;
  --forest:#1c3a30;
  --forest-2:#274c3f;
  --forest-3:#152a23;
  --ochre:#a4661f;
  --ochre-soft:#c98d45;
  --accent:#a8392c;
  --cream:#f1e8d6;
  --gold:#d3ab6a;
  --serif:"Noto Serif SC","Source Han Serif SC","Songti SC","STSong",SimSun,Georgia,"Times New Roman",serif;
  --sans:"Noto Sans SC","PingFang SC","Hiragino Sans GB","Microsoft YaHei","Segoe UI",system-ui,sans-serif;
  --shadow-s:0 1px 2px rgba(24,33,28,.05);
  --shadow-m:0 18px 40px -22px rgba(24,33,28,.35);
  --shadow-l:0 40px 90px -40px rgba(10,18,14,.55);
  --ease:cubic-bezier(.22,.61,.36,1);
}

*,*::before,*::after{box-sizing:border-box}
html{scroll-behavior:smooth;-webkit-text-size-adjust:100%;text-size-adjust:100%}
body{
  margin:0;
  background:var(--paper);
  color:var(--ink);
  font-family:var(--sans);
  font-size:16px;
  line-height:1.8;
  -webkit-font-smoothing:antialiased;
  text-rendering:optimizeLegibility;
}
body::before{
  content:"";
  position:fixed;inset:0;z-index:0;pointer-events:none;
  background:
    radial-gradient(circle at 12% 18%,rgba(201,141,69,.07),transparent 42%),
    radial-gradient(circle at 88% 72%,rgba(28,58,48,.06),transparent 46%),
    repeating-linear-gradient(0deg,rgba(24,33,28,.018) 0 1px,transparent 1px 3px);
  opacity:.9;
}
body.is-locked{overflow:hidden}
img{max-width:100%;display:block}
a{color:inherit;text-decoration:none}
button{font:inherit;color:inherit;background:none;border:0;cursor:pointer}
em{font-style:italic}
::selection{background:var(--forest);color:var(--cream)}

.wrap{width:min(1280px,100% - 64px);margin-inline:auto;position:relative;z-index:1}
.skip-link{position:absolute;left:-9999px;top:0;background:var(--forest);color:var(--cream);padding:10px 18px;z-index:200}
.skip-link:focus{left:12px;top:12px}

/* ---------- header ------------------------------------------------------- */
.site-header{
  position:sticky;top:0;z-index:60;
  backdrop-filter:blur(14px) saturate(120%);
  background:rgba(243,238,226,.86);
  border-bottom:1px solid rgba(24,33,28,.1);
}
.header-inner{display:flex;align-items:center;justify-content:space-between;height:72px}
.brand{display:flex;align-items:center;gap:12px}
.brand-mark{width:38px;height:38px;color:var(--forest);flex:none}
.brand-text{display:flex;flex-direction:column;line-height:1.1}
.brand-text strong{font-family:var(--serif);font-size:17px;letter-spacing:.12em;font-weight:600}
.brand-text em{font-style:normal;font-size:9.5px;letter-spacing:.32em;text-transform:uppercase;color:var(--muted)}
.site-nav{display:flex;gap:32px}
.site-nav a{
  font-size:13px;letter-spacing:.08em;color:var(--ink-2);position:relative;padding:6px 0;
}
.site-nav a::after{
  content:"";position:absolute;left:0;right:100%;bottom:0;height:1px;background:var(--forest);
  transition:right .45s var(--ease);
}
.site-nav a:hover::after,.site-nav a.is-active::after{right:0}
.site-nav a.is-active{color:var(--forest)}

/* ---------- hero --------------------------------------------------------- */
.hero{
  position:relative;min-height:min(88vh,820px);display:flex;align-items:flex-end;
  background:var(--forest-3);color:var(--cream);overflow:hidden;isolation:isolate;
  padding:120px 0 54px;
}
.hero-media{position:absolute;inset:0;z-index:-2}
.hero-media img{width:100%;height:100%;object-fit:cover;object-position:center 42%;filter:saturate(.92) contrast(1.03)}
.hero-veil{
  position:absolute;inset:0;z-index:-1;
  background:
    linear-gradient(180deg,rgba(12,22,18,.42) 0%,rgba(12,22,18,.18) 32%,rgba(12,22,18,.72) 78%,rgba(12,22,18,.94) 100%),
    linear-gradient(90deg,rgba(12,22,18,.72) 0%,rgba(12,22,18,.2) 58%,rgba(12,22,18,.05) 100%);
}
.hero-inner{width:min(1280px,100% - 64px)}
.eyebrow{
  display:flex;align-items:center;gap:14px;margin:0 0 22px;
  font-size:10.5px;letter-spacing:.34em;text-transform:uppercase;color:rgba(241,232,214,.78);
}
.eyebrow-line{display:block;width:44px;height:1px;background:var(--gold);flex:none}
.hero h1{
  margin:0;font-family:var(--serif);font-weight:500;
  font-size:clamp(4.2rem,9.5vw,8.2rem);line-height:.92;letter-spacing:.06em;
  text-shadow:0 20px 60px rgba(0,0,0,.35);
}
.hero h1 span{
  display:block;margin-top:18px;font-family:var(--sans);font-size:clamp(.8rem,1.4vw,1rem);
  font-weight:400;letter-spacing:.52em;text-transform:uppercase;color:var(--gold);text-shadow:none;
}
.hero-sub{
  max-width:540px;margin:26px 0 0;font-size:15.5px;line-height:2;color:rgba(241,232,214,.84);
  text-wrap:pretty;
}
.hero-stats{
  display:grid;grid-template-columns:repeat(4,minmax(0,1fr));gap:24px;
  margin-top:44px;padding-top:24px;border-top:1px solid rgba(241,232,214,.22);max-width:860px;
}
.hero-stats b{display:block;font-family:var(--serif);font-size:2rem;font-weight:400;line-height:1;letter-spacing:.02em}
.hero-stats span{display:block;margin-top:8px;font-size:10.5px;letter-spacing:.16em;text-transform:uppercase;color:rgba(241,232,214,.58)}
.hero-caption{
  position:absolute;right:max(32px,calc((100vw - 1280px)/2));bottom:18px;margin:0;
  font-size:10.5px;letter-spacing:.06em;color:rgba(241,232,214,.5);max-width:420px;text-align:right;
}

/* ---------- sections ----------------------------------------------------- */
.section{padding:104px 0;position:relative;scroll-margin-top:86px}
.section + .section{border-top:1px solid var(--line-soft)}
.section-head{max-width:760px;margin-bottom:56px}
.section-kicker{
  margin:0 0 14px;font-size:10.5px;letter-spacing:.3em;text-transform:uppercase;color:var(--ochre);
}
.section-head h2{
  margin:0;font-family:var(--serif);font-weight:500;font-size:clamp(2rem,3.6vw,2.9rem);
  letter-spacing:.08em;line-height:1.2;
}
.section-lead{margin:18px 0 0;font-size:16px;line-height:1.95;color:var(--muted);text-wrap:pretty}

/* ---------- overview ----------------------------------------------------- */
.overview-grid{display:grid;grid-template-columns:1.35fr .9fr;gap:64px;align-items:start}
.prose p{margin:0 0 22px;font-size:15.5px;line-height:2.05;color:var(--ink-2);text-wrap:pretty}
.prose p:last-child{margin-bottom:0}
.prose strong{font-weight:600;color:var(--forest);box-shadow:inset 0 -.5em 0 rgba(211,171,106,.22)}
.prose em{font-style:italic}
.fact-panel{
  display:grid;grid-template-columns:1fr 1fr;background:var(--paper-2);
  border:1px solid var(--line);box-shadow:var(--shadow-s);
}
.fact{padding:26px 24px 24px;border-right:1px solid var(--line-soft);border-bottom:1px solid var(--line-soft)}
.fact:nth-child(2n){border-right:0}
.fact:nth-child(n+3){border-bottom:0}
.fact-label{display:block;font-size:10px;letter-spacing:.24em;text-transform:uppercase;color:var(--muted-2)}
.fact b{display:block;margin:8px 0 4px;font-family:var(--serif);font-weight:500;font-size:2.1rem;line-height:1.1;color:var(--forest)}
.fact b small{font-size:.55em;margin-left:3px;color:var(--muted)}
.fact-note{display:block;font-size:11.5px;color:var(--muted);line-height:1.6}
.diagnosis{list-style:none;display:grid;grid-template-columns:repeat(4,1fr);gap:40px;margin:72px 0 0;padding:0}
.diagnosis li{border-top:2px solid var(--forest);padding-top:22px}
.diag-num{font-family:var(--serif);font-size:12px;letter-spacing:.2em;color:var(--ochre)}
.diagnosis h3{margin:10px 0 8px;font-family:var(--serif);font-size:19px;font-weight:600;letter-spacing:.06em}
.diagnosis p{margin:0;font-size:13.5px;line-height:1.9;color:var(--muted)}

/* ---------- tree --------------------------------------------------------- */
.tree-section{background:var(--forest-3);color:var(--cream);border-top:0}
.tree-section::before{
  content:"";position:absolute;inset:0;pointer-events:none;opacity:.5;
  background:
    radial-gradient(ellipse at 20% 0%,rgba(211,171,106,.12),transparent 55%),
    radial-gradient(ellipse at 90% 90%,rgba(60,110,90,.22),transparent 55%);
}
.tree-section .section-kicker{color:var(--gold)}
.tree-section .section-lead{color:rgba(241,232,214,.62)}
.tree-meta{display:flex;justify-content:space-between;align-items:flex-end;gap:28px;margin-bottom:26px;flex-wrap:wrap}
.rank-path{margin:0;font-size:11.5px;letter-spacing:.04em;color:rgba(241,232,214,.48);line-height:1.9}
.rank-path i{margin:0 8px;font-style:normal;opacity:.5}
.rank-path strong{color:rgba(241,232,214,.86);font-weight:500}
.tree-tools{display:flex;gap:10px}
.ghost-btn{
  padding:8px 16px;border:1px solid rgba(241,232,214,.26);border-radius:2px;
  font-size:12px;letter-spacing:.08em;color:rgba(241,232,214,.82);
  transition:background .3s,border-color .3s,color .3s;
}
.ghost-btn:hover{background:rgba(241,232,214,.1);border-color:rgba(241,232,214,.5)}
.tree-frame{
  border:1px solid rgba(241,232,214,.14);background:rgba(241,232,214,.025);
  box-shadow:inset 0 0 60px rgba(0,0,0,.25);
}
.tree-scroll{overflow-x:auto;overflow-y:hidden;padding:26px 30px 34px;scrollbar-width:thin;scrollbar-color:rgba(241,232,214,.25) transparent}
.tree-scroll::-webkit-scrollbar{width:9px;height:9px}
.tree-scroll::-webkit-scrollbar-thumb{background:rgba(241,232,214,.22);border-radius:9px}
.tree-scroll::-webkit-scrollbar-track{background:transparent}
#tree-svg{display:block;min-width:780px}
.tree-empty{padding:60px 30px;text-align:center;font-size:14px;color:rgba(241,232,214,.55)}
.tree-legend{
  display:flex;align-items:center;gap:34px;flex-wrap:wrap;margin:0 0 14px;
  font-size:12px;color:rgba(241,232,214,.62);
}
.tree-legend em{font-style:italic;opacity:.85}
.dot{display:inline-block;width:8px;height:8px;border-radius:50%;margin-right:9px;vertical-align:1px}
.dot.channa{background:#b9cdb9}
.dot.parachanna{background:#e0a15c}
.legend-hint{margin-left:auto;font-size:11px;color:rgba(241,232,214,.38);letter-spacing:.04em}
.tree-hint{display:none;margin:0 0 10px;font-size:11px;color:rgba(241,232,214,.45);letter-spacing:.04em}
.tree-note{
  margin:30px 0 0;padding-top:18px;border-top:1px solid rgba(241,232,214,.12);
  max-width:900px;font-size:12.5px;line-height:1.95;color:rgba(241,232,214,.5);
}

/* tree drawing */
.tree-link{fill:none;stroke:rgba(241,232,214,.28);stroke-width:1}
.tree-link.genus{stroke-width:1.4}
.tree-link.is-dim{opacity:.25}
.tree-link.is-hot{stroke:rgba(211,171,106,.95);stroke-width:1.8}
.tree-node{cursor:pointer}
.tree-node circle{fill:#15251f;stroke-width:1.4}
.tree-node.genus circle{stroke-width:1.6}
.tree-node:hover circle{fill:var(--gold)}
.tree-zh{font-family:var(--serif);font-size:14px;fill:rgba(241,232,214,.92);letter-spacing:.04em}
.tree-sci{font-family:var(--serif);font-style:italic;font-size:12px;fill:rgba(241,232,214,.55)}
.tree-size{font-family:var(--sans);font-size:10.5px;fill:rgba(241,232,214,.35)}
.tree-root-zh{font-family:var(--serif);font-size:22px;fill:var(--cream);letter-spacing:.14em}
.tree-root-sci{font-family:var(--sans);font-size:10px;fill:rgba(241,232,214,.45);letter-spacing:.3em;text-transform:uppercase}
.tree-genus-zh{font-family:var(--serif);font-size:17px;fill:var(--cream);letter-spacing:.1em}
.tree-genus-sci{font-family:var(--serif);font-style:italic;font-size:12.5px;fill:rgba(241,232,214,.55)}
.tree-count{font-family:var(--sans);font-size:10px;fill:rgba(241,232,214,.4);letter-spacing:.08em}
.tree-row.is-dim{opacity:.28}
.tree-row.is-hot .tree-zh{fill:var(--gold)}
.tree-row.is-hot .tree-sci{fill:rgba(211,171,106,.8)}
.tree-toggle{fill:rgba(241,232,214,.5);font-size:11px;font-family:var(--sans)}

/* ---------- filters ------------------------------------------------------ */
.filter-bar{border-top:1px solid var(--line);border-bottom:1px solid var(--line);padding:22px 0 20px;margin-bottom:26px}
.search-box{
  display:flex;align-items:center;gap:10px;border:1px solid var(--line);background:var(--paper-2);
  padding:0 14px;height:46px;max-width:420px;transition:border-color .3s,box-shadow .3s;
}
.search-box:focus-within{border-color:var(--forest);box-shadow:0 0 0 3px rgba(28,58,48,.08)}
.search-box svg{width:17px;height:17px;flex:none;fill:none;stroke:var(--muted);stroke-width:1.6;stroke-linecap:round}
.search-box input{
  flex:1;min-width:0;border:0;background:transparent;outline:0;font:inherit;font-size:14px;color:var(--ink);
  padding:2px 0;
}
.search-box input::placeholder{color:var(--muted-2)}
.search-clear{font-size:18px;line-height:1;color:var(--muted);padding:4px}
.filter-row{display:flex;align-items:center;gap:16px;margin-top:14px;flex-wrap:wrap}
.filter-row--end{margin-top:20px;justify-content:space-between}
.filter-label{font-size:11px;letter-spacing:.22em;text-transform:uppercase;color:var(--muted-2);width:34px;flex:none}
.chips{display:flex;gap:8px;flex-wrap:wrap}
.chip{
  border:1px solid var(--line);border-radius:999px;padding:5px 14px;font-size:12.5px;
  color:var(--ink-2);transition:all .25s var(--ease);line-height:1.7;background:rgba(250,246,236,.4);
}
.chip em{font-style:italic;font-size:.94em;opacity:.8;margin-left:2px}
.chip:hover{border-color:var(--forest);background:var(--paper-2)}
.chip.is-active{background:var(--forest);border-color:var(--forest);color:var(--cream)}
.chip-count{opacity:.55;margin-left:2px;font-size:11px}
.switch{display:flex;align-items:center;gap:10px;font-size:12.5px;color:var(--ink-2);cursor:pointer;user-select:none}
.switch input{position:absolute;opacity:0;width:0;height:0}
.switch-track{
  width:36px;height:20px;border-radius:20px;background:var(--line);position:relative;transition:background .3s;flex:none;
}
.switch-track::after{
  content:"";position:absolute;top:2px;left:2px;width:16px;height:16px;border-radius:50%;
  background:var(--paper-2);transition:transform .3s var(--ease);box-shadow:0 1px 2px rgba(0,0,0,.2);
}
.switch input:checked + .switch-track{background:var(--forest)}
.switch input:checked + .switch-track::after{transform:translateX(16px)}
.switch input:focus-visible + .switch-track{outline:2px solid var(--ochre);outline-offset:2px}
.sort-box{display:flex;align-items:center;gap:8px;font-size:12px;color:var(--muted)}
.sort-box select{
  font:inherit;font-size:12.5px;color:var(--ink-2);border:1px solid var(--line);background:var(--paper-2);
  padding:6px 10px;border-radius:2px;cursor:pointer;
}
.result-meta{margin:0 0 24px;font-size:12.5px;color:var(--muted);letter-spacing:.03em}
.result-meta b{font-weight:600;color:var(--ink);font-family:var(--serif);font-size:15px}
.result-meta span{margin-left:8px;color:var(--muted-2)}

/* ---------- cards -------------------------------------------------------- */
.cards{display:grid;grid-template-columns:repeat(auto-fill,minmax(228px,1fr));gap:24px}
.cards[hidden]{display:none}
.empty-state[hidden]{display:none}
.trade-block{margin-top:66px;padding-top:36px;border-top:1px solid var(--line)}
.trade-block[hidden]{display:none}
.trade-head{margin-bottom:24px}
.trade-badge{display:inline-block;margin-bottom:10px;padding:3px 9px;border:1px dashed var(--ochre);color:var(--ochre);font-size:10px;letter-spacing:.16em}
.trade-head h3{margin:0;font-family:var(--serif);font-size:22px;font-weight:500;letter-spacing:.04em;color:var(--ink)}
.trade-head h3 em{font-style:italic;font-size:15px;color:var(--muted);margin-left:2px}
.trade-head p{margin:7px 0 0;font-size:12.5px;line-height:1.8;color:var(--muted)}
.trade-tag{background:rgba(168,102,31,.92);color:var(--cream)}
.trade-cards{max-width:280px}
.card{
  display:flex;flex-direction:column;text-align:left;padding:0;background:var(--paper-2);
  border:1px solid var(--line-soft);position:relative;overflow:hidden;
  transition:transform .45s var(--ease),box-shadow .45s var(--ease),border-color .45s;
  opacity:0;transform:translateY(14px);
}
.card.is-in{opacity:1;transform:none}
.card:hover{transform:translateY(-5px);box-shadow:var(--shadow-m);border-color:var(--line)}
.card:focus-visible{outline:2px solid var(--ochre);outline-offset:3px}
.card-figure{display:block;position:relative;aspect-ratio:4/3;overflow:hidden;background:var(--paper-3)}
.card-figure img{width:100%;height:100%;object-fit:cover;transition:transform .8s var(--ease),filter .4s}
.card:hover .card-figure img{transform:scale(1.045)}
.card-genus{
  position:absolute;top:12px;left:12px;z-index:2;padding:3px 9px 4px;
  background:rgba(243,238,226,.92);backdrop-filter:blur(4px);
  font-size:9.5px;letter-spacing:.2em;text-transform:uppercase;color:var(--forest);
}
.card-genus.parachanna{color:var(--ochre)}
.card-tag{
  position:absolute;right:12px;bottom:12px;z-index:2;display:flex;align-items:center;gap:6px;
  padding:3px 9px;background:rgba(18,30,25,.72);backdrop-filter:blur(4px);color:var(--cream);
  font-size:10px;letter-spacing:.08em;
}
.card-tag i{width:6px;height:6px;border-radius:50%;background:currentColor}
.card-body{padding:17px 18px 18px;display:flex;flex-direction:column;gap:2px}
.card-zh{font-family:var(--serif);font-size:20px;line-height:1.35;letter-spacing:.04em;color:var(--ink)}
.card-zh .aster{font-size:12px;color:var(--ochre);vertical-align:super;margin-left:3px}
.card-zh .none{font-family:var(--sans);font-size:13px;color:var(--muted-2);letter-spacing:.02em}
.card-latin{font-family:var(--serif);font-style:italic;font-size:13px;color:var(--ink-2);line-height:1.5}
.card-meta{
  display:flex;align-items:center;gap:10px;margin-top:11px;padding-top:11px;border-top:1px solid var(--line-soft);
  font-size:11px;color:var(--muted);
}
.card-meta .size{font-variant-numeric:tabular-nums}
.card-meta .sep{width:1px;height:10px;background:var(--line)}
.iucn{display:inline-flex;align-items:center;gap:5px;letter-spacing:.04em}
.iucn i{width:6px;height:6px;border-radius:50%;background:var(--muted-2);display:inline-block}
.iucn.LC i{background:#5c8a5a}.iucn.NT i{background:#c2a13c}.iucn.VU i{background:#d07a35}
.iucn.EN i{background:#bf4a33}.iucn.CR i{background:#8f2a24}.iucn.DD i{background:#8b8f96}.iucn.NE i{background:#b3b0a4}
.card-photo-none{
  width:100%;height:100%;display:flex;flex-direction:column;align-items:center;justify-content:center;gap:10px;
  background:
    repeating-linear-gradient(45deg,rgba(24,33,28,.035) 0 1px,transparent 1px 9px),
    var(--paper-3);color:var(--muted);
}
.card-photo-none svg{width:96px;height:44px;opacity:.4}
.card-photo-none span{font-size:10px;letter-spacing:.18em;text-transform:uppercase}

/* ---------- about -------------------------------------------------------- */
.about{background:var(--paper-2)}
.about-grid{display:grid;grid-template-columns:1fr 1fr;gap:56px 64px}
.about-block h3{
  margin:0 0 14px;font-family:var(--serif);font-size:18px;font-weight:600;letter-spacing:.08em;
  padding-bottom:12px;border-bottom:1px solid var(--line);
}
.about-block p{margin:0 0 14px;font-size:14px;line-height:1.95;color:var(--ink-2)}
.about-block p:last-child{margin-bottom:0}
.about-block a{color:var(--forest);border-bottom:1px solid rgba(28,58,48,.3)}
.about-block a:hover{border-bottom-color:var(--forest)}
.refs{margin:0;padding-left:20px;font-size:13px;line-height:1.9;color:var(--muted)}
.refs li{margin-bottom:8px}
.refs em{font-style:italic}
.mark-trans{color:var(--ochre);font-weight:600}
.disclaimer{
  margin-top:56px;padding:22px 26px;border-left:3px solid var(--ochre);
  background:rgba(201,141,69,.08);font-size:13.5px;color:var(--ink-2);line-height:1.9;
}
.disclaimer p{margin:0}

/* ---------- footer ------------------------------------------------------- */
.site-footer{background:var(--forest-3);color:rgba(241,232,214,.62);padding:52px 0}
.footer-inner{display:flex;align-items:center;justify-content:space-between;gap:28px;flex-wrap:wrap}
.footer-brand{display:flex;align-items:center;gap:12px;color:var(--cream)}
.footer-brand svg{width:34px;height:34px;opacity:.85}
.footer-brand div{display:flex;flex-direction:column;line-height:1.3}
.footer-brand strong{font-family:var(--serif);font-size:16px;letter-spacing:.14em;font-weight:500}
.footer-brand span{font-size:9.5px;letter-spacing:.3em;text-transform:uppercase;color:rgba(241,232,214,.45)}
.footer-note{margin:0;font-size:12px;line-height:1.9}
.footer-note a{border-bottom:1px solid rgba(241,232,214,.3)}
.footer-note a:hover{color:var(--cream);border-color:var(--cream)}
.footer-top{font-size:12px;letter-spacing:.1em;border-bottom:1px solid rgba(241,232,214,.25);padding-bottom:2px}
.footer-top:hover{color:var(--cream);border-color:var(--cream)}

/* ---------- modal -------------------------------------------------------- */
.modal{position:fixed;inset:0;z-index:100;display:none}
.modal.is-open{display:block}
.modal-backdrop{position:absolute;inset:0;background:rgba(9,16,13,.72);backdrop-filter:blur(6px);animation:fade .35s both}
.modal-panel{
  position:absolute;inset:50% auto auto 50%;transform:translate(-50%,-50%);
  width:min(1160px,calc(100vw - 64px));height:min(86vh,820px);max-height:min(86vh,820px);
  background:var(--paper);box-shadow:var(--shadow-l);
  display:grid;grid-template-columns:46% 54%;overflow:hidden;
  animation:rise .45s var(--ease) both;
}
@keyframes fade{from{opacity:0}to{opacity:1}}
@keyframes rise{from{opacity:0;transform:translate(-50%,-47%)}to{opacity:1;transform:translate(-50%,-50%)}}
.modal-close{
  position:absolute;top:14px;right:14px;z-index:5;width:40px;height:40px;border-radius:50%;
  background:rgba(243,238,226,.92);color:var(--ink);font-size:24px;line-height:1;
  display:grid;place-items:center;box-shadow:var(--shadow-s);transition:background .25s,transform .25s;
}
.modal-close:hover{background:var(--cream);transform:rotate(90deg)}
.modal-arrow{
  position:absolute;top:14px;z-index:5;width:40px;height:40px;border-radius:50%;
  background:rgba(243,238,226,.92);color:var(--ink);box-shadow:var(--shadow-s);
  display:grid;place-items:center;font-size:17px;transition:background .25s;
}
.modal-arrow:hover{background:var(--cream)}
.modal-arrow.prev{right:108px}
.modal-arrow.next{right:62px}
.modal-media{position:relative;background:#0e1a16;min-height:100%;overflow:hidden}
.modal-media img{position:absolute;inset:18px;width:calc(100% - 36px);height:calc(100% - 36px);object-fit:contain}
.modal-credit{
  position:absolute;left:0;right:0;bottom:0;margin:0;padding:36px 20px 16px;
  font-size:10px;line-height:1.6;color:rgba(241,232,214,.72);
  background:linear-gradient(180deg,transparent,rgba(10,18,14,.85));
}
.modal-credit a{color:rgba(241,232,214,.92);border-bottom:1px solid rgba(241,232,214,.35)}
.modal-body{padding:46px 44px 40px;overflow-y:auto;scrollbar-width:thin}
.modal-kicker{display:flex;align-items:center;gap:10px;font-size:10px;letter-spacing:.24em;text-transform:uppercase;color:var(--ochre);margin:0 0 12px}
.modal-kicker i{width:24px;height:1px;background:currentColor;display:block}
#modal-title{margin:0;font-family:var(--serif);font-size:34px;font-weight:500;line-height:1.25;letter-spacing:.03em}
.modal-latin{font-family:var(--serif);font-style:italic;font-size:17px;color:var(--ink-2);margin:6px 0 0}
.modal-author{font-size:11.5px;color:var(--muted);margin:2px 0 0}
.modal-en{margin:14px 0 0;font-size:13px;color:var(--muted)}
.modal-path{
  margin:22px 0 0;padding:12px 14px;background:rgba(28,58,48,.05);border-left:2px solid var(--forest);
  font-size:11.5px;line-height:1.8;color:var(--ink-2);letter-spacing:.02em;
}
.modal-path em{font-style:italic}
.modal-facts{display:grid;grid-template-columns:1fr 1fr;gap:0;margin:24px 0 0;border-top:1px solid var(--line)}
.modal-fact{padding:13px 0;border-bottom:1px solid var(--line-soft)}
.modal-fact:nth-child(2n){padding-left:22px;border-left:1px solid var(--line-soft)}
.modal-fact.wide{grid-column:1 / -1;padding-left:0;border-left:0}
.modal-fact dt{font-size:9.5px;letter-spacing:.2em;text-transform:uppercase;color:var(--muted-2);margin:0 0 3px}
.modal-fact dd{margin:0;font-size:13.5px;color:var(--ink);line-height:1.65}
.modal-fact dd em{font-style:italic}
.modal-section{margin:26px 0 0}
.modal-section h3{font-family:var(--serif);font-size:12px;letter-spacing:.24em;text-transform:uppercase;color:var(--forest);margin:0 0 10px;padding-bottom:8px;border-bottom:1px solid var(--line)}
.modal-section p{margin:0;font-size:13.5px;line-height:1.95;color:var(--ink-2)}
.morph-list{display:flex;flex-wrap:wrap;gap:8px;margin:0;padding:0;list-style:none}
.morph-list li{font-size:12px;padding:5px 10px;background:var(--paper-2);border:1px solid var(--line-soft);color:var(--ink-2)}
.morph-list b{font-weight:600;color:var(--forest);margin-left:4px;font-variant-numeric:tabular-nums}
.modal-links{display:flex;gap:18px;flex-wrap:wrap;margin-top:26px;padding-top:18px;border-top:1px solid var(--line);font-size:11.5px}
.modal-links a{border-bottom:1px solid rgba(28,58,48,.35);color:var(--forest);padding-bottom:1px}
.modal-links a:hover{border-color:var(--forest)}
.iucn-badge{display:inline-flex;align-items:center;gap:6px;font-size:12.5px}
.iucn-badge i{width:7px;height:7px;border-radius:50%;background:#b3b0a4}
.iucn-badge.LC i{background:#5c8a5a}.iucn-badge.NT i{background:#c2a13c}.iucn-badge.VU i{background:#d07a35}
.iucn-badge.EN i{background:#bf4a33}.iucn-badge.CR i{background:#8f2a24}.iucn-badge.DD i{background:#8b8f96}
.empty-state{text-align:center;padding:70px 20px;color:var(--muted);font-size:14px}
.link-btn{color:var(--forest);border-bottom:1px solid currentColor;margin-left:8px;font-size:13px}

/* ---------- reveal ------------------------------------------------------- */
.reveal{opacity:0;transform:translateY(16px);transition:opacity .8s var(--ease),transform .8s var(--ease)}
.reveal.is-in{opacity:1;transform:none}

/* ---------- responsive --------------------------------------------------- */
@media (max-width:1080px){
  .overview-grid{grid-template-columns:1fr;gap:40px}
  .diagnosis{grid-template-columns:1fr 1fr;gap:32px}
  .about-grid{grid-template-columns:1fr;gap:40px}
  .modal-panel{grid-template-columns:42% 58%}
  .hero-stats{grid-template-columns:repeat(4,1fr);gap:16px}
}
@media (max-width:820px){
  .wrap{width:calc(100% - 40px)}
  .site-nav{display:flex;gap:12px;flex-wrap:nowrap}
  .site-nav a{font-size:11.5px;letter-spacing:.01em;white-space:nowrap}
  .brand-text em{display:none}
  .brand-text strong{font-size:15px;letter-spacing:.06em}
  .brand-mark{width:30px;height:30px}
  .header-inner{height:62px;gap:12px}
  .hero{min-height:78vh;padding-bottom:40px}
  .hero-stats{grid-template-columns:1fr 1fr;gap:18px 20px}
  .hero-stats b{font-size:1.6rem}
  .hero-caption{display:none}
  .section{padding:72px 0}
  .section-head{margin-bottom:36px}
  .diagnosis{grid-template-columns:1fr;gap:24px}
  .modal-panel{grid-template-columns:1fr;height:92vh;max-height:92vh;width:calc(100vw - 28px)}
  .modal-media{min-height:34vh;max-height:34vh}
  .modal-body{padding:28px 24px 32px;max-height:58vh}
  #modal-title{font-size:26px}
  .modal-arrow{top:12px}
  .modal-arrow.prev{right:106px}
  .modal-arrow.next{right:60px}
  .legend-hint{display:none}
  .tree-hint{display:block}
  .tree-scroll{padding:18px 16px 24px}
  .cards{grid-template-columns:repeat(auto-fill,minmax(210px,1fr));gap:18px}
  .footer-inner{flex-direction:column;align-items:flex-start;gap:18px}
}
@media (max-width:520px){
  .hero h1{letter-spacing:.02em}
  .hero-sub{font-size:14px}
  .card-zh{font-size:18px}
  .filter-label{width:100%}
  .cards{grid-template-columns:1fr 1fr;gap:12px}
  .card-body{padding:12px 12px 14px}
  .card-zh{font-size:16px}
  .card-latin{font-size:11px}
  .card-meta{font-size:10px;gap:6px}
}
@media (prefers-reduced-motion:reduce){
  *,*::before,*::after{animation-duration:.001ms !important;transition-duration:.001ms !important}
  html{scroll-behavior:auto}
  .card{opacity:1;transform:none}
}

/* ---------- print -------------------------------------------------------- */
@media print{
  .site-header,.modal,.filter-bar,.tree-tools,.hero-caption{display:none}
  .hero{min-height:auto;background:#fff;color:#000}
  .hero-media,.hero-veil{display:none}
  .section{padding:20px 0;break-inside:avoid}
  .cards{grid-template-columns:repeat(3,1fr)}
  .card{opacity:1;transform:none;break-inside:avoid}
}